Tlayúa Formation

Tlayúa Formation
Stratigraphic range: Albian PreꞒ Ꞓ O S D C P T J K Pg N
Type	Geological formation
Sub-units	Lower, Middle and Upper Members
Underlies	Unconformity with Eocene-Oligocene Pie de Vaca Formation
Overlies	Orizaba Formation
Thickness	Lower: 50 m Middle: 35 m Upper: 40 m
Lithology
Primary	Limestone
Other	Mudstone
Location
Coordinates	Tlayúa Formation (Mexico)

The Tlayúa Formation is an Early Cretaceous (late Albian) geological formation near Tepexi de Rodríguez, Puebla.^[1]

Paleobiota

The formation contains a diverse array of vertebrate and invertebrate fossils. About 70% of the macrofossils are osteichthyan fish.^[2] Other vertebrates include chelonians, pterosaurs, lepidosaurs, and crocodiles.^[3] Cyanobacteria, foraminifera, algae, gymnosperms, sponges, cnidarians, annelids, gastropods, ammonites, bivalves, arachnids, insects, isopods, anomurans, brachyurans, crinoids, echinoids, holothuroids, stelleroids, and ophiuroids, have also been recovered from the Tlayúa Formation.^[3]^[4]^[5]

Lepidosaurs

Genus	Species	Notes
Huehuecuetzpalli	H. mixtecus	A primitive lizard
Pamizinsaurus	P. tlayuaensis	An osteoderm-covered sphenodontian
Ankylosphenodon^[6]	A. pachyostosus	An aquatic sphenodontian
Tepexisaurus^[7]	T. tepexii	A basal scincomorph

Other vertebrates

Mainly after González-Rodríguez (2016)^[8]

Araripichthys webri
Armigatus carrenoae
Armigatus felixi
Axelrodichthys cf. araripensis
Cipactlichthys scutatus
Epaelops martinezi
Macrosemiocotzus americanus
Michin csernai
Notagogus novomundi
Nusaviichthys nerivelai
Pachyamia mexicana
Paraclupea seilacheri
Quetzalichthys perrilliatae
Ranulfoichthys dorsonudum
Teoichthys brevipina
Teoichthys kallistos
Tepexichthys aranguthyrom
Tlalocbatos applegatei
Tlayuamichin itztli
Unamichthys espinosai
Ellimmichthys sp.
cf. Amblysemius
cf. Bananogmius
Belonostomus sp.
cf. Brannerion
Lepidotes sp.
cf. Lycoptera
cf. Megalops
Neoproscinetes sp.
Notelops sp.
cf. Paraelops
cf. Rhacolepis
Vinctifer sp.
cf. Yabrudichthys
Gonorhynchiform indet.
Testudines indet.

Arthropods

Archaeoniscus aranguthyorum
Atocatle ranulfoi
Ixtahua benjamini
Protaegla miniscula
Tepexicarcinus tlayuaensis
Sphaeromatidae sp.

Echinoderms

Ophiactis applegatei
Paleopentacta alencasterae
Parapsolus tlayuensis
Astropecten sp.
Echinaster sp.
Ophidiaster sp.
Plutonaster sp.
Tamaria sp.

Plants

References

^ Applegate, S. (1988). "A new genus and species of a holostean belonging to the family Ophiopsidae, Teoichthys kallistos, from the Cretaceous, near Tepexi de Rodriguez, Puebla" (PDF). Revista Mexicana de Ciencias Geológicas. 7 (2): 200–205.
^ Alvarado-Ortega J, González-Rodríguez KA, Blaco-Piñón A, Espinosa-Arrubarrena L, Ovalles-Damián E (2006). "Mesozoic Osteichthyans of Mexico". In Vega, FJ, TG Nyborg, MC Perrilliat, M. Montellano-Ballesteros, SRS Cevallos-Ferriz. (eds.). Studies on Mexican Paleontology, Topics on Geobiology 24. Dordrecht, Netherlands: Springer Verlag. pp. 169–207. ISBN 1402039859.
^ ^a ^b Vega FJ, Bruce NL, Serrano L, Bishop GA, Perrilliat MD (2005). "A review of the Lower Cretaceous (Tlayúa Formation: Albian) Crustacea from Tepexi de Rodríguez, Puebla, Central Mexico" (PDF). Bulletin of the Mizunami Fossil Museum (32): 25–30. Archived from the original (PDF) on 2011-06-12.
^ Buitrón-Sánchez, Blanca Estela; Durán-Gónzalez, Alicia; Martín-Cao-Romero, Carolina; Solís-Marín, Francisco Alonso; Laguarda-Figueras, Alfredo. "Lower Cretaceous (Albian) Asteroidea (Echinodermata) from Tepexi de Rodriguez, Puebla, Mexico". Revista de Biología Tropical. 63 (2): 7–15.
^ Solé, Jesús; Pi-Puig, Teresa; Bermúdez-Chávez, Cynthia; Garduño-Martínez, Diana; Alvarado-Ortega, Jesús (2022-12-05). "Mineralogy, geochemistry, and K-Ar dating of feldspars and clays from an exceptional Cretaceous fossil locality (Tlayúa, Puebla, Mexico): Insights into the depositional and diagenetic ages and processes". Chemical Geology. 612: 121134. doi:10.1016/j.chemgeo.2022.121134. ISSN 0009-2541.
^ Reynoso, V. H. (2000). "An Unusual Aquatic Sphenodontian (Reptilia: Diapsida) from the Tlayua Formation (Albian), Central Mexico". Journal of Paleontology. 74 (1): 133–148. doi:10.1017/S0022336000031310. JSTOR 1306891.
^ Vega, Francisco J.; Nyborg, Torrey G.; María del Carmen Perrilliat; Marisol Montellano-Ballesteros; Sergio R.S Cevallos-Ferriz; Sara A. Quiroz-Barroso (2006). "Research on Fossil Amphibians and Reptiles". In Landman, Neil H. (ed.). Studies on Mexican Paleontology. Vol. 24. Douglas S. Jones. Dordrecht, Netherlands: Springer. p. 214. ISBN 1-4020-3882-8. Retrieved 31 May 2011.
^ Rodríguez, Katia González; Fielitz, Christopher. "CRETACEOUS OSTEICHTHYAN FISH ASSEMBLAGES FROM MEXICO". {{cite journal}}: Cite journal requires |journal= (help)